Vespiform

Vespiforms, also called Giant Wasps, were an ancient and wise insectoid species, physically described as giant wasps by Donna Noble. They had hives in the Silfrax Galaxy. They were a hench-species of the Quarks before they turned on them.

Biology
Vespiforms were a species of insectoids which resembled oversized wasps. Adults were about three metres tall. They were powerful creatures, with large stingers which could be regrown within hours of use.

Vespiforms could change into other species. The transformations were accompanied by a glowing, purple-pink haze and left behind morphic residue.

When in another form, Vespiforms were genetically compatible with the mimicked species. They could interbreed with members of any other species. Any offspring would be locked in one form until put under stress, when they would turn into their Vespiform shape. (TV: The Unicorn and the Wasp)

Technology
Vespiforms had telepathic recorders. These devices held the essence of the Vespiform cultural heritage, and served as a sort of "instant school". The entirety of what it meant to be a Vespiform could be transferred from the recorder to the mind in a matter of seconds. The device also transferred the thoughts of any being in very close proximity to it as well. Were the thoughts of a non-Vespiform to blend with the information contained in the recorder, the results could be unpredictable. (TV: The Unicorn and the Wasp)

History
The Quarks attempted to use Giant Wasps to attack the Second Doctor and Jamie McCrimmon on Gano, but the creatures attacked the Quarks instead. (COMIC: The Killer Wasps)

Some time prior to Earth's 19th century, the Vespiforms developed intergalactic space travel. This allowed at least one of the species to travel from the Silfrax Galaxy to Mutter's Spiral. At least one Vespiform went to Earth. (WC: )

In 1885, a Vespiform (simply known as Christopher) met Clemency Eddison in India and had a child with her. He gave her a large, purple crystal set in a necklace, which she called "the Firestone". When he soon thereafter drowned during a monsoon, the Firestone became her only memory of him, and she wore it often.

Some months later, Clemency gave up their child for adoption; the morés of the time frowned on unwed parents. The child became a reverend, Arnold Golightly, a vicar in the parish that included the Eddison's estate. At the age of forty in 1926, he discovered his Vespiform nature. Because of his proximity to the Firestone — in reality a Vespiform telepathic recorder — his Vespiform cultural heritage was transferred to him from the device. The transfer also included some of its wearer's thoughts. Lady Eddison was reading an Agatha Christie novel at the time of the transfer, so Golightly absorbed some of the aspects of the characters in the novel. Golightly started committing various murders in the style of an Agatha Christie novel. The Tenth Doctor, Donna Noble and Agatha Christie herself investigated his crimes, and forced Golightly to publicly reveal his Vespiform nature. Ultimately, Donna killed him by throwing the Firestone into the lake. Golightly followed and drowned, but relieved Agatha of his connection with her in his dying moment. This caused the confusion and amnesia that marked the author's famous disappearance. (TV: The Unicorn and the Wasp)

The Sixth Doctor and Mel had a "harrowing escapade" with the Quarks and the Giant Wasps on Earth, in 1999. (PROSE: Millennial Rites) In a parallel universe created by the Quantum Archangel, there remained a Quark processor unit from a "run-in" with the Quarks and Giant Wasps". Mel later felt that she wanted her life back from before she had met the Doctor, and she mentally listed some of the aliens she had encounted, one pair being "the Quarks and their giant wasp servants". Tom Dering, the President of the United States in 2010 had an encounter with the Quarks and the Giant Wasps in Central Park. (PROSE: The Quantum Archangel)

Long after the events at Eddison Manor, the Firestone was dredged up from the lake and sold at auction to Jack Harkness. In the 21st century, he claimed to be in possession of the only Vespiform telepathic recorder known to exist on Earth.

Jack Harkness had heard whispers of a war between Quark rebels and Vespiforms, but he could not confirm these stories. (WC: )

Behind the scenes

 * According to The Brilliant Book 2012, a book that contains non-narrative based information, Amy Pond and Rory Williams might have encountered some Vespiform in the Gardens of Zul-Thep in the year 3104. (REF: The Brilliant Book 2012)
 * Russell T Davies' original concept for the Shadow Proclamation scene in The Stolen Earth included an appearance by several buzzing Vespiform, but his plans to feature a "rogues gallery" of alien races in the scene fell through; ultimately the Judoon were the only returning race featured.
 * While the Vespiforms and Giant Wasps were physically identical, it was only WC: Monster File: Vespiform that drew an overt link showing the species to be one and the same.